Charles Darwin

Charles Robert Darwin was born on 12 February 1809 at his family's home, The Mount, in Shrewsbury, Shropshire. He was the fifth of six children of wealthy society doctor and financier Robert Darwin and Susannah Darwin (née Wedgwood). His grandfathers Erasmus Darwin and Josiah Wedgwood were both prominent abolitionists. Erasmus Darwin had praised general concepts of evolution and common descent in his Zoonomia (1794), a poetic fantasy of gradual creation including undeveloped ideas anticipating concepts his grandson expanded.

Both families were largely Unitarian, though the Wedgwoods were adopting Anglicanism. Robert Darwin, a freethinker, had baby Charles baptised in November 1809 in the Anglican St Chad's Church, Shrewsbury, but Charles and his siblings attended the local Unitarian Church with their mother. The eight-year-old Charles already had a taste for natural history and collecting when he joined the day school run by its preacher in 1817. That July, his mother died. From September 1818, he joined his older brother Erasmus in attending the nearby Anglican Shrewsbury School as a boarder.

Darwin spent the summer of 1825 as an apprentice doctor, helping his father treat impoverished people in Shropshire, before going to the well-regarded University of Edinburgh Medical School with his brother Erasmus in October 1825. Darwin found lectures dull and surgery distressing, so he neglected his studies. He learned taxidermy in around 40 daily hour-long sessions from John Edmonstone, a Black Briton from Demerara in the South American rainforest, who had been taught there by Charles Waterton, and when brought to Scotland was freed from slavery.

In Darwin's second year at the university, he joined the Plinian Society, a student natural-history group featuring lively debates in which radical democratic students with materialistic views challenged orthodox religious concepts of science. He assisted Robert Edmond Grant's investigations of the anatomy and life cycle of marine invertebrates in the Firth of Forth, and on 27 March 1827 presented at the Plinian his own discovery that black spores found in oyster shells were the eggs of a skate leech.

One day, Grant praised Lamarck's evolutionary ideas. Darwin was astonished by Grant's audacity, but had recently read similar ideas in his grandfather Erasmus' journals. Darwin was rather bored by Robert Jameson's natural-history course, which covered geology – including the debate between neptunism and plutonism. He learned the classification of plants. He assisted with work on the collections of the University Museum, one of the largest museums in Europe at the time.

Darwin's neglect of medical studies annoyed his father, who sent him to Christ's College, Cambridge in January 1828 to study for a Bachelor of Arts degree as the first step towards becoming an Anglican country parson. Darwin was unqualified for Cambridge's Tripos exams and was required instead to join the ordinary degree course. He preferred riding and shooting to studying. At Cambridge, Darwin joined a society known as the "Glutton Club," which consumed "birds and beasts which were before unknown to human palate." Darwin would continue to eat the meat of exotic animals for the rest of his life; on the voyage of the Beagle, he ate puma, iguanas, giant tortoises, armadillos, and mistakenly ate a rhea which he intended to study.

During the first few months of Darwin's enrolment at Christ's College, his second cousin William Darwin Fox was still studying there. Fox impressed him with his butterfly collection, introducing Darwin to entomology and influencing him to pursue beetle collecting. He did this zealously and had some of his finds published in James Francis Stephens' Illustrations of British entomology (1829–1932).

Through Fox, Darwin became a close friend and follower of botany professor John Stevens Henslow. He met other leading parson-naturalists who saw scientific work as religious natural theology, becoming known to these dons as "the man who walks with Henslow". When his own exams drew near, Darwin applied himself to his studies and was delighted by the language and logic of William Paley's Evidences of Christianity (1795). In his final examination in January 1831, Darwin did well, coming tenth out of 178 candidates for the ordinary degree.

Darwin had to stay at Cambridge until June 1831. He studied Paley's Natural Theology or Evidences of the Existence and Attributes of the Deity (first published in 1802), which made an argument for divine design in nature, explaining adaptation as God acting through laws of nature. He read John Herschel's new book, Preliminary Discourse on the Study of Natural Philosophy (1831), which described the highest aim of natural philosophy as understanding such laws through inductive reasoning based on observation, and Alexander von Humboldt's Personal Narrative of scientific travels in 1799–1804. Inspired with "a burning zeal" to contribute, Darwin planned to visit Tenerife with some classmates after graduation to study natural history in the tropics. In preparation, he joined Adam Sedgwick's geology course, then on 4 August travelled with him to spend a fortnight mapping strata in Wales.

After leaving Sedgwick in Wales, Darwin spent a few days with student friends at Barmouth. He returned home on 29 August to find a letter from Henslow proposing him as a suitable (if unfinished) naturalist for a self-funded supernumerary place on HMS Beagle with captain Robert FitzRoy, a position for a gentleman rather than "a mere collector". The ship was to leave in four weeks on an expedition to chart the coastline of South America. Robert Darwin objected to his son's planned two-year voyage, regarding it as a waste of time, but was persuaded by his brother-in-law, Josiah Wedgwood II, to agree to (and fund) his son's participation. Darwin took care to remain in a private capacity to retain control over his collection, intending it for a major scientific institution.

After delays, the voyage began on 27 December 1831; it lasted almost five years. As FitzRoy had intended, Darwin spent most of that time on land investigating geology and making natural history collections, while HMS Beagle surveyed and charted coasts. He kept careful notes of his observations and theoretical speculations. At intervals during the voyage, his specimens were sent to Cambridge together with letters including a copy of his journal for his family. He had some expertise in geology, beetle collecting and dissecting marine invertebrates, but in all other areas, was a novice and ably collected specimens for expert appraisal. Despite suffering badly from seasickness, Darwin wrote copious notes while on board the ship. Most of his zoology notes are about marine invertebrates, starting with plankton collected during a calm spell.

On their first stop ashore at St Jago in Cape Verde, Darwin found that a white band high in the volcanic rock cliffs included seashells. FitzRoy had given him the first volume of Charles Lyell's Principles of Geology, which set out uniformitarian concepts of land slowly rising or falling over immense periods,[II] and Darwin saw things Lyell's way, theorising and thinking of writing a book on geology. When they reached Brazil, Darwin was delighted by the tropical forest, but detested the sight of slavery there, and disputed this issue with FitzRoy.

The survey continued to the south in Patagonia. They stopped at Bahía Blanca, and in cliffs near Punta Alta, Darwin made a significant find of fossil bones of huge extinct mammals beside modern seashells, indicating recent extinction with no signs of change in climate or catastrophe. He found bony plates like a giant version of the armour on local armadillos. From a jaw and tooth, he identified the gigantic Megatherium, then from Cuvier's description thought the armour was from this animal. The finds were shipped to England, and scientists found the fossils of great interest.

On rides with gauchos into the interior to explore geology and collect more fossils, Darwin gained social, political, and anthropological insights into both native and colonial people at a time of revolution, and learnt that two types of rhea had separate but overlapping territories. Further south, he saw stepped plains of shingle and seashells as raised beaches at a series of elevations. He read Lyell's second volume and accepted its description of "centres of creation" of species, but his discoveries and theorising challenged Lyell's ideas of smooth continuity and of extinction of species.

In Tierra del Fuego, Darwin formed the incorrect belief that the archipelago was devoid of reptiles.

Three Fuegians on board, who had been seized during the first Beagle voyage and then given Christian education in England, were returning with a missionary. Darwin found them friendly and civilised, yet at Tierra del Fuego he met "miserable, degraded savages", as different as wild from domesticated animals. He remained convinced that, despite this diversity, all humans were interrelated with a shared origin and potential for improvement towards civilisation. Unlike his scientist friends, he now thought there was no unbridgeable gap between humans and animals. A year on, the mission had been abandoned. The Fuegian they had named Jemmy Button lived like the other natives, had a wife, and had no wish to return to England.

Darwin experienced an earthquake in Chile in 1835 and saw signs that the land had just been raised, including mussel beds stranded above high tide. High in the Andes, he saw seashells and several fossil trees that had grown on a sand beach. He theorised that as the land rose, oceanic islands sank, and coral reefs around them grew to form atolls.

On the geologically new Galápagos Islands, Darwin looked for evidence attaching wildlife to an older "centre of creation". He found mockingbirds allied to those in Chile but differing from island to island. He heard that slight variations in the shape of tortoise shells showed which island they came from, but failed to collect them, even after eating tortoises taken on board as food. In Australia, the marsupial rat-kangaroo and the platypus seemed so unusual that Darwin thought it was almost as though two distinct Creators had been at work. He found the Aboriginal Australians "good-humoured & pleasant", their numbers depleted by European settlement.

FitzRoy investigated how the atolls of the Cocos (Keeling) Islands had formed. The survey supported Darwin's theorising. FitzRoy began writing the official Narrative of the Beagle voyages, and after reading Darwin's diary, he proposed incorporating it into the account. Darwin's Journal was eventually rewritten as a separate third volume, on geology and natural history.

In Cape Town, South Africa, Darwin and FitzRoy met John Herschel, who had recently written to Lyell praising his uniformitarianism as opening bold speculation on "that mystery of mysteries, the replacement of extinct species by others" as "a natural in contradistinction to a miraculous process".

When organising his notes as the ship sailed home, Darwin wrote that, if his growing suspicions about the mockingbirds, the tortoises and the Falkland Islands fox were correct, "such facts undermine the stability of Species", then cautiously added "would" before "undermine". He later wrote that such facts "seemed to me to throw some light on the origin of species".

Without Darwin's knowledge, extracts from his letters to Henslow had been read to scientific societies, printed as a pamphlet for private distribution among members of the Cambridge Philosophical Society, and reported in magazines, including The Athenaeum. Darwin first heard of this at Cape Town, and at Ascension Island read of Sedgwick's prediction that Darwin "will have a great name among the Naturalists of Europe".

On 2 October 1836, Beagle anchored at Falmouth, Cornwall.

The Darwins had ten children: two died in infancy, and Annie's death at the age of ten had a devastating effect on her parents. Charles was a devoted father and uncommonly attentive to his children. Whenever they fell ill, he feared that they might have inherited weaknesses from inbreeding due to the close family ties he shared with his wife and cousin, Emma Wedgwood. He examined inbreeding in his writings, contrasting it with the advantages of outcrossing in many species.

Charles Waring Darwin, born in December 1856, was the tenth and last of the children. Emma Darwin was aged 48 at the time of the birth, and the child was mentally subnormal and never learnt to walk or talk. He probably had Down syndrome, which had not then been medically described. The evidence is a photograph by William Erasmus Darwin of the infant and his mother, showing a characteristic head shape, and the family's observations of the child. Charles Waring died of scarlet fever on 28 June 1858, when Darwin wrote in his journal: "Poor dear Baby died."

Of his surviving children, George, Francis and Horace became Fellows of the Royal Society, distinguished as an astronomer, botanist and civil engineer, respectively. All three were knighted. Another son, Leonard, went on to be a soldier, politician, economist, eugenicist, and mentor of the statistician and evolutionary biologist Ronald Fisher.

Darwin's family tradition was nonconformist Unitarianism, while his father and grandfather were freethinkers, and his baptism and boarding school were Church of England. When going to Cambridge to become an Anglican clergyman, he did not "in the least doubt the strict and literal truth of every word in the Bible". He learned John Herschel's science which, like William Paley's natural theology, sought explanations in laws of nature rather than miracles and saw adaptation of species as evidence of design. On board HMS Beagle, Darwin was quite orthodox and would quote the Bible as an authority on morality. He looked for "centres of creation" to explain distribution, and suggested that the very similar antlions found in Australia and England were evidence of a divine hand.

Upon his return, he expressed a critical view of the Bible's historical accuracy and questioned the basis for considering one religion more valid than another. In the next few years, while intensively speculating on geology and the transmutation of species, he gave much thought to religion and openly discussed this with his wife Emma, whose beliefs similarly came from intensive study and questioning.

The theodicy of Paley and Thomas Malthus vindicated evils such as starvation as a result of a benevolent creator's laws, which had an overall good effect. To Darwin, natural selection produced the good of adaptation but removed the need for design. He was increasingly troubled by the problem of evil and he could not see the work of an omnipotent deity in all the pain and suffering, such as the ichneumon wasp paralysing caterpillars as live food for its eggs. Though he thought of religion as a tribal survival strategy, in a letter to Asa Gray in 1860, Darwin also confessed that he could not "anyhow be contented to view this wonderful universe (as) the result of brute force," making him reluctant to give up the idea of God as an ultimate lawgiver.

Darwin remained close friends with the vicar of Downe, John Brodie Innes, and continued to play a leading part in the parish work of the church, but from c. 1849 would go for a walk on Sundays while his family attended church. He considered it "absurd to doubt that a man might be an ardent theist and an evolutionist" and, though reticent about his religious views, in a letter to John Fordyce in 1879, he wrote that "I have never been an atheist in the sense of denying the existence of a God. – I think that generally ... an agnostic would be the most correct description of my state of mind".

At other times he believed in a First Cause: This follows from the extreme difficulty, or rather the impossibility, of conceiving this immense and wonderful universe, including man with his capacity for looking far backwards and far into futurity, as the result of blind chance or necessity. When thus reflecting I feel compelled to look to a First Cause having an intelligent mind in some degree analogous to that of man; and I deserve to be called a Theist. This conclusion was strong in my mind about the time, as far as I can remember, when I wrote the Origin of Species; and it is since that time that it has very gradually with many fluctuations become weaker. [...] I cannot pretend to throw the least light on such abstruse problems. The mystery of the beginning of all things is insoluble by us; and I for one must be content to remain an Agnostic.

The "Lady Hope Story", published in 1915, claimed that Darwin had reverted to Christianity on his sickbed. The claims were repudiated by Darwin's children and have been dismissed as false by historians.

Darwin's views on social and political issues reflected his time and social position. He grew up in a family of Whig reformers who, like his uncle Josiah Wedgwood, supported electoral reform and the emancipation of slaves. Darwin was passionately opposed to slavery.

Taking taxidermy lessons in 1826 from the freed slave John Edmonstone, whom Darwin long recalled as "a very pleasant and intelligent man", reinforced his belief that black people shared the same feelings, and could be as intelligent as people of other races. He took the same attitude to native people he met on the Beagle voyage. Though commonplace in Britain at the time, Silliman and Bachman noticed the contrast with slave-owning America. Around twenty years later, racism became a feature of British society, but Darwin remained strongly against slavery, against "ranking the so-called races of man as distinct species", and against ill-treatment of native people.[VII]

Darwin's interaction with Yaghans (Fuegians) such as Jemmy Button during the second voyage of HMS Beagle had a profound impact on his view of indigenous peoples. At his arrival in Tierra del Fuego, he made a colourful description of "Fuegian savages". This view changed as he came to know the Yaghan people more in detail. By studying the Yaghans, Darwin concluded that a number of basic emotions by different human groups were the same and that mental capabilities were roughly the same as for Europeans. While interested in Yaghan culture, Darwin failed to appreciate their deep ecological knowledge and elaborate cosmology until the 1850s when he inspected a dictionary of Yaghan detailing 32,000 words. He saw that European colonisation would often lead to the extinction of native civilisations, and "tr[ied] to integrate colonialism into an evolutionary history of civilization analogous to natural history".

Darwin's view of women was that men's eminence over them was the outcome of sexual selection, a view disputed by Antoinette Brown Blackwell in her 1875 book The Sexes Throughout Nature.

Darwin was intrigued by his half-cousin Francis Galton's argument, introduced in 1865, that statistical analysis of heredity showed that moral and mental human traits could be inherited, and principles of animal breeding could apply to humans. In The Descent of Man, Darwin noted that aiding the weak to survive and have families could lose the benefits of natural selection, but cautioned that withholding such aid would endanger the instinct of sympathy, "the noblest part of our nature", and factors such as education could be more important. When Galton suggested that publishing research could encourage intermarriage within a "caste" of "those who are naturally gifted", Darwin foresaw practical difficulties and thought it "the sole feasible, yet I fear utopian, plan of procedure in improving the human race", preferring to simply publicise the importance of inheritance and leave decisions to individuals. Francis Galton named this field of study "eugenics" in 1883,[VIII] after Darwin's death, and his theories were cited to promote eugenic policies.

Darwin's fame and popularity led to his name being associated with ideas and movements that, at times, had only an indirect relation to his writings, and sometimes went directly against his express comments.

Thomas Malthus had argued that population growth beyond resources was ordained by God to get humans to work productively and show restraint in getting families; this was used in the 1830s to justify workhouses and laissez-faire economics. Evolution was by then seen as having social implications, and Herbert Spencer's 1851 book Social Statics based ideas of human freedom and individual liberties on his Lamarckian evolutionary theory.

Soon after the Origin was published in 1859, critics derided his description of a struggle for existence as a Malthusian justification for the English industrial capitalism of the time. The term Darwinism was used for the evolutionary ideas of others, including Spencer's "survival of the fittest" as free-market progress, and Ernst Haeckel's polygenistic ideas of human development. Writers used natural selection to argue for various, often contradictory, ideologies such as laissez-faire dog-eat-dog capitalism, colonialism, and imperialism. However, Darwin's holistic view of nature included "dependence of one being on another"; thus pacifists, socialists, liberal social reformers and anarchists such as Peter Kropotkin stressed the value of cooperation over struggle within a species. Darwin himself insisted that social policy should not simply be guided by concepts of struggle and selection in nature.

After the 1880s, the eugenics movement developed on ideas of biological inheritance, and for scientific justification of their ideas appealed to some concepts of Darwinism. In Britain, most shared Darwin's cautious views on voluntary improvement and sought to encourage those with good traits in "positive eugenics". During the "Eclipse of Darwinism", a scientific foundation for eugenics was provided by Mendelian genetics. Negative eugenics to remove the "feebleminded" was widely popular across the political spectrum in the United States, Canada, and Australia. Belief in negative eugenics led to the introduction of compulsory sterilisation laws in the United States, followed by several other countries. Subsequently, Nazi eugenics brought the field into disrepute.[VII]

The term "Social Darwinism" was used infrequently from around the 1890s, but became popular as a derogatory term in the 1940s when used by Richard Hofstadter to attack the laissez-faire conservatism of those like William Graham Sumner who opposed reform and socialism. Since then, it has been used as a term of abuse by those opposed to what they think are the moral consequences of evolution.

Darwin was a prolific writer. Even without the publication of his works on evolution, he would have had a considerable reputation as the author of The Voyage of the Beagle, as a geologist who had published extensively on South America and had solved the puzzle of the formation of coral atolls, and as a biologist who had published the definitive work on barnacles. While On the Origin of Species dominates perceptions of his work, The Descent of Man and The Expression of the Emotions in Man and Animals had considerable impact, and his books on plants including The Power of Movement in Plants were innovative studies of great importance, as was his final work on The Formation of Vegetable Mould Through the Action of Worms.

As Alfred Russel Wallace put it, Darwin had "wrought a greater revolution in human thought within a quarter of a century than any man of our time – or perhaps any time", having "given us a new conception of the world of life, and a theory which is itself a powerful instrument of research; has shown us how to combine into one consistent whole the facts accumulated by all the separate classes of workers, and has thereby revolutionised the whole study of nature". The paleoanthropologist Trenton Holliday states that "Darwin is rightly considered to be the preeminent evolutionary scientist of all time". Ernst Mayr considered On the Origin of Species to be, second to the Bible, the second most important book in history with regards to its influence on human thought, and he considered the scientific revolution that Darwin's theory of evolution ushered in to be "perhaps the most fundamental of all the intellectual revolutions in the history of mankind".

By around 1880, most scientists were convinced of evolution as descent with modification, though few agreed with Darwin that natural selection "has been the main but not the exclusive means of modification". During "the eclipse of Darwinism" scientists explored alternative mechanisms. Then Ronald Fisher incorporated Mendelian genetics in The Genetical Theory of Natural Selection, leading to population genetics and the modern evolutionary synthesis, which continues to develop. Scientific discoveries have confirmed and validated Darwin's key insights. The biologist Theodosius Dobzhansky said that "Nothing in biology makes sense except in the light of evolution."

Geographical features given his name include Darwin Sound and Mount Darwin, both named while he was on the Beagle voyage, and Darwin Harbour, named by his former shipmates on its next voyage, which eventually became the location of Darwin, the capital city of Australia's Northern Territory. Darwin's name was given, formally or informally, to numerous plants and animals, including many he had collected on the voyage.

In 1908, the Linnean Society of London began awards of the Darwin–Wallace Medal, to mark fifty years from the joint reading on 1 July 1858 of papers by Darwin and Wallace publishing their theory. Further awards were made in 1958 and 2008; since 2010, the awards have been annual. Darwin College, a postgraduate college at Cambridge University founded in 1964, is named after the Darwin family. From 2000 to 2017, UK£10 banknotes issued by the Bank of England featured Darwin's portrait printed on the reverse, along with a hummingbird and HMS Beagle. Darwin's bicentennial was celebrated with a series of postage stamps in the United Kingdom.

The Smithsonian National Museum of Natural History has a bronze statue of Charles Darwin in its Deep Time Hall, which features Darwin seated on a bench with a notebook containing his "tree of life" sketch. The statue was sculpted by David Clendining and was installed as the centrepiece of the hall, which focuses on Darwinian evolution.